Publikasjoner
NIBIOs ansatte publiserer flere hundre vitenskapelige artikler og forskningsrapporter hvert år. Her finner du referanser og lenker til publikasjoner og andre forsknings- og formidlingsaktiviteter. Samlingen oppdateres løpende med både nytt og historisk materiale. For mer informasjon om NIBIOs publikasjoner, besøk NIBIOs bibliotek.
2011
Sammendrag
In 2005 an extensive test program including field tests was set up in order to obtain more data on the durability and long term performance of modified wood and semi-durable wood species. One of the main challenges for modified wood is to predict accurate service life time in UC3 (Use use class 3, above ground) and UC4 (use class 4, in soil or fresh water contact). So far, data from in-service conditions are rare, while several studies have evaluated the durability in lab or field test exposure. However, there is still a lack of studies comparing replicate modified wood products in both field and lab exposure. This study evaluates the efficacy of modified wood in AWPA E10, three different types of soil in lab (ENV 807), three test fields in-ground (EN 252) and two close to ground (horizontal double layer test) set-ups at two test sites. The test material includes furfurylated, acetylated and thermally modified wood in addition to reference treated and control samples. In laboratory, both furfurylated, acetylated and thermally modified pine (212ºC) performed well. The modified wood samples performed at the same level, or better, than the reference CC and CCA preservatives in retentions for UC4 applications. In the horizontal double layer test, five years is still too short time to be able to draw firm conclusions. However, in the most accelerated HDL set-up, all controls have failed or are moderately to severely decayed whereas most preservative treated, furfurylated and acetylated wood are sound or only slightly decayed. After 5 years of testing CCA-preserved wood performs better in-ground in field tests than in lab tests, whereas modified wood generally performs slightly poorer. Just like in the lab tests, however, acetylated wood performs equal to CCA-preserved wood in UC4. Furfurylated wood performs equal to or better than UC3 level preservative treated wood. Thermally modified wood actually performs much poorer than all preservative treated wood references. Finally, natural durability classification of the same treatment in different lab and field tests was surprisingly similar.
Forfattere
Gry AlfredsenSammendrag
Modified wood can provide protection against a range of wood deteriorating organisms. Several hypotheses have been put forward for the mode of action against wood decaying fungi, including inhibition of action of specific enzymes, but they still need further testing. This paper summarizes results from a project focusing on molecular studies of fungal colonization in modified wood. The focus has been on furfurylated wood, but also thermally modified and acetylated wood has been studied. Among the main finding was that wood modifications have an effect on the exploitation face of both brown and white rot colonization, but not on the exploration face. As already reported in a range of papers wood modification effects the wood moisture content, and this was confirmed within this project. New information was gathered about the effect on gene expression. Even before any mass loss was detected, differences in gene expression were measured. [...]
Sammendrag
Brown rot is the most common and destructive type of fungal decay for wood in service. These fungi depolymerize preferentially the structural carbohydrates, cellulose and hemicellulose in the cell wall leaving oxidized lignin behind. Modified wood can provide protection against a variety of wood deteriorating organisms, including decay fungi. However, there is still little known about the mode of function of the different wood modifications concerning the decay resistance. The biochemical mechanisms and gene products induced in brown rot during growth in modified wood are poorly understood. In this paper the data collected from mass loss studies and qPCR and qRT-PCR were used for profiling growth dynamics and gene expression of the brown rot fungus Postia placenta in different wood substrates through different stages of decay. Pinus sylvestris (L.) sapwood was used for the following treatments and modifications: chromated copper arsenate CCA (0.67%), furfurylation (WPG 37), thermal modification (D212) and acetylation (WPG 23). Untreated Pinus sylvestris (L.) sapwood was used as control. Samples were taken at different time intervals from 2 to 26 weeks. The highest mass loss and the highest fungal DNA content were found in the control samples while acetylated wood had the lowest mass loss and fungal DNA content. These results reflect a close relation of mass loss and fungal DNA content, both reflecting the amount of Postia placenta decaying the samples. Generally, expression of the investigated genes was highest in CCA treated wood. In the beginning of the incubation of all treated wood samples, the genes coding for oxidative metabolic activity had higher expression levels than the untreated control. In the end of the incubation most of these genes were less expressed than in the untreated control. The genes used for carbohydrate metabolism and the alcohol oxidase showed a significant decrease after 14 weeks of incubation. At the same time an increase in gene expression of an enzyme putative involved in lignin decomposition was detected.
Sammendrag
Some of the most common Norwegian wood species were tested in a Double layer test in South East Norway. After eight years of exposure the highest decay rating (≥3) was found in Scots pine sapwood, Norway spruce, alder, birch and aspen. Two wood types had decay rate ≤1: Scots pine heartwood and cedar. Wood moisture was logged and compared with precipitation during a two month period the second year of exposure. Scots pine sapwood had higher wood moisture content than Norway spruce, and a good correlation was found between precipitation and wood moisture content. When comparing similar materials exposed at three different geographical locations in Southern Norway, the samples exposed in Bergen had higher decay rating than samples exposed at Ås and Oslo.
Sammendrag
Moisture is often recognised as a key factor regarding the long time performance of wooden products, and one of the main challenges for timber products is to predict accurate service life in use class 3 (not covered above ground) and use class 4 (in soil or fresh water contact). A range of durability classification studies have been performed both in field and laboratory. But for several wood species information regarding the durability in use class 3 is lacking. Also, there is still a lack of studies comparing replicate wood products in different field exposure situations. This study evaluates the natural durability of different North European wood species in two different climates and in two different use classes. The wood species were compared with imported species and two preservative treatments. The overall picture shows a higher decay rating for wood species tested in ground contact compared with the results from the above ground “Double layer tests”. Moreover, the woods tested in Western Norway are more decayed than those tested in Eastern Norway. These findings can be explained by higher decay risk in use class 4 than in use class 3, and higher decay risk in a humid climate (Western Norway) than in a dry climate (Eastern Norway). The results indicate similar ranking of the durability of the wood species regardless of the environment they have been exposed to. The results from a linear regression show that MOE-loss of the mini-stakes after three years describes 70 % of the variation in decay rating of the “Double layer” stakes after six years exposure in Western Norway. This result strongly indicates that MOE-loss can be a prospective tool for rapid field testing of natural durability of wood.
Sammendrag
To understand the defence mechanisms utilized by decay fungi when exposed to different wood protection systems the study of gene expression can give us some answers. When the DNA sequences are known, primers can be designed to detect transcripts of genes with gene products related to basic cellular processes and hyphal growth. The characteristic gene products induced in different fungi by different wood protection systems can be identified. Studies on the expression of fungal genes will give us a better understanding of the fungal degradation of wood and we can optimize wood protection systems. Hence, no single technique will give us the answer to all questions about the decay of wood we need to gather small pieces of the puzzle using different approaches. The aim of the present study was to investigate the effects of acetylation level on the growth of Postia placenta with regard to amount of total DNA and gene expression targeting 7 different genes. This paper presents preliminary results after 4 weeks of incubation. The results presented in this paper are parts of a larger project which reaches over a period of 36 weeks with sampling times after 12, 20, 28 and 36 weeks. We found no mass loss in the acetylated samples after 4 weeks of incubation in a modified soil-block test. The presence of P. placenta DNA and the absence of mass loss could indicate on an inability of the mycelia to establish a wood exploitation phase. Two genes related to carbohydrate metabolism were expressed in a higher amount in P. placenta during growth on untreated wood than during growth on acetylated wood. However, for a third gene, also related to carbohydrate metabolism, the relationship was the opposite. Two genes related to oxidative metabolism were expressed in a higher amount in P. placenta during growth on acetylated wood than during growth on untreated wood and another two genes related to oxidative metabolism showed inconsistent results.
Sammendrag
De siste 30 årene har halvparten av gammelskogen på Varaldskogen blitt snauhogd. Likevel har storfuglen klart seg ganske bra, mens orrfuglen har gått litt ned.
Sammendrag
Along the succession gradient of the boreal forest ecosystem, black grouse Tetrao tetrix inhabits the early and capercaillie Tetrao urogallus the latest stages. When converting old forest to clearcuts and plantations, commercial forestry has therefore been assumed to affect capercaillie negatively and to be favourable to black grouse. During a 30-year period (1979–2008) we monitored sympatric populations of the two species in a forest in southeast Norway based on annual spring and autumn censuses and radio-marked birds. During this period, the proportion of old, semi-natural forest was halved and clearcuts and young plantations increased accordingly. The grouse populations did not change as predicted. While the trend in August numbers of adult black grouse declined, males more than females, abundance of adult capercaillie remained unchanged. Number of males at leks showed similar patterns. Equally surprising, breeding success (number of chicks per female in August) of both species increased, thus indicating that the populations were regulated more by variation in adult survivorship than by recruitment of young birds. No correlations were found with changing climatic factors (precipitation and temperatures in winter and spring, snow depth and time of snow melt), except that year-to-year breeding success was positively correlated with minimum temperatures during 2 weeks posthatch. The results are explained by a combination of more flexible habitat selection than previously assumed and a changing predator regime: In the early period, nearly all capercaillie leks were located in old, semi-natural forest, but as plantations grew older (>30 years), new leks were established there. Similarly, while young capercaillie broods used old semi-natural forest almost exclusively when the study started, they frequently used middle-aged plantations, especially those with a ground cover of bilberry Vaccinium myrtillus, when these became common in later years. The increasing breeding success could largely be explained by more females rearing chicks successfully, presumably due to a marked decline in the main nest predator, the red fox Vulpes vulpes. A practice of thinning of the old, semi-natural forest some years prior to final harvesting probably facilitated predation of black grouse by goshawks Accipiter gentilis. Contrary to many beliefs, our results indicate that both capercaillie and black grouse are quite tolerant to changes in forest management regimes. In our study, numerical and functional responses of predators (mainly red fox and goshawk) apparently played a more important role in regulating grouse numbers than habitat factors per se.
Sammendrag
The redlisted epiphytic lichen Evernia divaricata has only rarely been found with fruiting or sorediate thalli, and never so in Norway. August 2009, we reinvented a previously known locality within the Trillemarka-Rollagsfjell nature reserve and found, for the first time in Norway, fertile thalli abundantly along a 1.2 km part of a small brook. Closer examination of samples taken from the same locality in 2003 revealed that immature fruiting-bodies (apothecia) were present on a few thalli at that time. Today the site is characterized by old mixed conifer forest and small bogs. Two additional Norwegian localities have been found to harbor fertile E. divaricata. At yet another locality we found one thallus with abundant large soralia, also recorded for the first time in Norway. Lack of small sprouting thalli with basal holdfasts suggests that dispersal, up till now, mainly have occurred by means of thallus fragmentation, thereby hampering efficient long-distance dispersal.
Sammendrag
Skogsfuglforskningen på Varaldskogen startet opp i 1979 etter en periode med svært lave bestander over hele landet. Etter 25 år med intensivt bestandsskogbruk mente mange at flatehogsten var hovedårsaken til bestandsnedgangen. På Varaldskogen var allerede halvparten av den gamle plukkhogde skogen avvirket, spesielt hadde det gått hardt utover de frodige granliene før staten tok over eiendommen etter Billerud i 1968. Mye av terrenget bestod av grasrike flater og tett ungskog i h.kl. 2. Tiurleikene og skogsfuglkullene befant seg i den gamle skogen. Reven hadde gode tider med mye markmus på hogstflatene, og den livnærte seg av egg og kyllinger når det ikke var museår...