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NIBIOs ansatte publiserer flere hundre vitenskapelige artikler og forskningsrapporter hvert år. Her finner du referanser og lenker til publikasjoner og andre forsknings- og formidlingsaktiviteter. Samlingen oppdateres løpende med både nytt og historisk materiale. For mer informasjon om NIBIOs publikasjoner, besøk NIBIOs bibliotek.

2010

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The furfurylation process is an extensively investigated wood modification process. Furfuryl alcohol molecules penetrate into the wood cell wall and polymerize in situ. This results in a permanent swelling of the wood cell walls. It is unclear whether or not chemical bonds exist between the furfuryl alcohol polymer and the wood. In the present study, five different wood species were used, both hardwoods and softwoods. They were treated with three different furfurylation procedures and leached according to three different leaching methods. The present study shows that, in general, the leachates from furfurylated wood have low toxicity. It also shows that the choice of leaching method is decisive for the outcome of the toxicity results. Earlier studies have shown that leachates from wood treated with furfuryl alcohol prepolymers have higher toxicity to Vibrio fischeri than leachates from wood treated with furfuryl alcohol monomers. This is probably attributable to differences in leaching of chemical compounds. The present study shows that this difference in the toxicity most likely cannot be attributed to maleic acid, furan, furfural, furfuryl alcohol, or 2-furoic acid. However, the difference might be caused by the two substances 5-hydroxymethylfurfural and 2,5-furandimethanol. The present study found no difference in the amount of leached furfuryl alcohol between leachates from furfurylated softwood and furfurylated hardwood species. Earlier studies have indicated differences in grafting of furfuryl alcohol to lignin. However, nothing was found in the present study that could support this. The leachates of furfurylated wood still need to be investigated further to identify the chemical differences between wood furfurylated with furfuryl alcohol monomers and furfuryl alcohol prepolymers.

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Constitutive and inducible terpene production is involved in conifer resistance against bark beetles and their associated fungi. In this study 72 Norway spruce (Picea abies) were randomly assigned to methyl jasmonate (MJ) application, inoculation with the bluestain fungus Ceratocystis polonica, or no-treatment control. We investigated terpene levels in the stem bark of the trees before treatment, 30 days and one year after treatment using GC–MS and two-dimensional GC (2D-GC) with a chiral column, and monitored landing and attack rates of the spruce bark beetle, Ips typographus, on the trees by sticky traps and visual inspection. Thirty days after fungal inoculation the absolute amount and relative proportion of (+)-3-carene, sabinene, and terpinolene increased and (+)-α-pinene decreased. Spraying the stems with MJ tended to generally increase the concentration of most major terpenes with minor alteration to their relative proportions, but significant increases were only observed for (−)-β-pinene and (−)-limonene. Fungal inoculation significantly increased the enantiomeric ratio of (−)-α-pinene and (−)-limonene 1 month after treatment, whereas MJ only increased that of (−)-limonene. One year after treatment, both MJ and fungal inoculation increased the concentration of most terpenes relative to undisturbed control trees, with significant changes in (−)-β-pinene, (−)-β-phellandrene and some other compounds. Terpene levels did not change in untreated stem sections after treatment, and chemical induction by MJ and C. polonica thus seemed to be restricted to the treated stem section. The enantiomeric ratio of (−)-α-pinene was significantly higher and the relative proportions of (−)-limonene were significantly lower in trees that were attractive to bark beetles compared to unattractive trees. One month after fungal inoculation, the total amount of diterpenes was significantly higher in putative resistant trees with shorter lesion lengths than in putative susceptible trees with longer lesions. Thus, terpene composition in the stem bark may be related to resistance of Norway spruce against I. typographus and C. polonica.

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Norwegian agriculture has, as in most western-European countries, gone through several periods of change during the last 100 years. Pronounced changes have occurred in production systems and the spatial organisation of farm land, as well as agricultural policy. During the last 50 years, official statistics document a marked decline in the number of active farms. This decline has caused concern, as Norway traditionally has had an agricultural policy that emphasises self-sufficiency and rural settlement. Yet statistics also show that the amount of agricultural land in use has remained the same. This is usually explained through a larger proportion of tenanted land, as technological progress has allowed production levels to be maintained with a smaller workforce. Studies elsewhere in Europe show, however, that tenancy may not promote the same levels of investment and landscape management as owner occupation. To assess the potential impact of this change on Norway's landscape (and its value as both a cultural and tourism resource) we analyse tenancy patterns in Norwegian agriculture between 1999 and 2003. In particular we note that, even if owner occupation remains strong nationally, when the statistics are broken down by municipality, tenancy has increased significantly in some areas. This has left large areas of land managed as tenancies by a relatively small number of farmers, including parts of the iconic west coast fjords. We conclude therefore that further work is urgently required to establish whether the effects of tenancy seen elsewhere apply to Norway, whether this exposes key landscapes to increased risk of abandonment and if so what appropriate political responses there could be.

Sammendrag

Ophiostoma spp. include important pathogens of trees and causal agents of sapstain. These fungi infect wounds on trees and are typically carried by insects, especially bark beetles. Ophiostoma spp. on coniferous hosts in the Northern Hemisphere are well-known. However, other than for the serious pathogens O. ulmi and O. novo-ulmi, very little research has been done on the occurrence of this group of fungi on native broad-leaved trees, especially in the Nordic countries. In this study, surveys were conducted in several areas of Norway to isolate Ophiostoma spp. associated with wounds on native broad-leaved trees belonging to the genera Betula, Fagus, Quercus, Sorbus and Tilia. Morphological studies and comparisons of DNA sequences for the ITS, 5.8S and part of the beta-tubulin gene regions were used to confirm the identity of the fungi collected. Ophiostoma spp., and especially their Pesotum anamorphs, were common on wounds on the trees sampled. In most cases, they were associated with wood stain. Ophiostoma spp. collected included predominantly O. quercus, O. borealis sp. nov., and O. denticiliatum. The results of this study emphasize that the diversity of Ophiostoma spp. on broad-leaved trees is still incompletely understood in Norway and other European countries.

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This study investigated the feasibility of extracting and chipping hardwood crowns for energy after motor-manual thinning in stands of common beech. Large crowns were extracted and chipped from stands where only sawlogs had been produced, while small crowns were extracted and chipped from stands where sawlogs and firewood had been harvested. The fuel chip yield was 15 m3 solid ha-1 when extracting and chipping large crowns, while it was 8 m3 solid ha-1 when extracting and chipping small crowns. The productivity for extracting and chipping large crowns was 8.5 m3 solid per workplace hour, and for small crowns was 5.9 m3 solid per workplace hour. Extracting and chipping large crowns gave a net income of €167 ha-1 (€11 m-3 solid), while extracting and chipping small crowns gave a lower net income of €23 ha-1 (€3 m-3 solid). The study showed that extracting and chipping large hardwood crowns is feasible and can make a substantial contribution to woody biomass feedstocks. Four product-mix alternatives were considered, but the marginal differences in outcome led the authors to recommend that in addition to sawlogs only one product, firewood or chips, should be produced in each stand.

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Riparian forests (RF) growing along streams, rivers and lakes comprise more than 2% of the forest area in the Nordic countries (considering a 10 m wide zone from the water body). They have special ecological functions in the landscape. They receive water and nutrients from the upslope areas, are important habitats for biodiversity, have large soil carbon stores, but may emit more greenhouse gases (GHG) than the uplands. In this article, we present a review of the environmental services related to water protection, terrestrial biodiversity, carbon storage and greenhouse gas dynamics provided by RF in the Nordic countries. We discuss the benefits and trade-offs when leaving the RF as a buffer against the impacts from upland forest management, in particular the impacts of clear cutting. Forest buffers are effective in protecting water quality and aquatic life, and have positive effects on terrestrial biodiversity, particularly when broader than 40 m, whereas the effect on the greenhouse gas exchange is unclear.

Sammendrag

In this study, the efficiency of a small multi-tree felling head, mounted on a farm tractor with a timber trailer was studied, when harvesting small trees for energy in thinnings. Both separate loading and direct loading of the felled trees was studied. Time studies were carried out in a mixed stand of Norway spruce (Picea abies (L.) Karst) and birch (Betula pubescens Ehrh.). The time consumption of the work elements in the different work methods was formulated by regression analysis, where the independent variables were tree size and degree of accumulation. The average size of the harvested trees was 0.035 m3. The time consumption for the harvesting and loading were similar for the two studied methods, 20 minutes per m3 at a tree size of 0.035 m3, but the two methods showed different characteristics for different tree sizes and level of accumulation. The direct loading method had the highest productivity when more than 0.1 m3 were collected in the felling cycle, whereas the separate loading method had the highest productivity when less than 0.05 m3 were collected in the felling cycle. The total effective time consumption for harvesting and forwarding the biomass 300 meters to roadside landing was 27 minutes per m3. The efficiency of the initial felling and collecting of the small trees was the main challenge. Both the harvesting technique and harvesting technology needs further development to provide a feasible production chain for woodfuel from energy thinning.

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Terrestrial biosphere models are indispensable tools for analyzing the biosphere-atmosphere exchange of carbon and water. Evaluation of these models using site level observations scrutinizes our current understanding of biospheric responses to meteorological variables. Here we propose a novel model-data comparison strategy considering that CO2 and H2O exchanges fluctuate on a wide range of timescales. Decomposing simulated and observed time series into subsignals allows to quantify model performance as a function of frequency, and to localize model-data disagreement in time. This approach is illustrated using site level predictions from two models of different complexity, Organizing Carbon and Hydrology in Dynamic Ecosystems (ORCHIDEE) and Lund-Potsdam-Jena (LPJ), at four eddy covariance towers in different climates. Frequency-dependent errors reveal substantial model-data disagreement in seasonal-annual and high-frequency net CO2 fluxes. By localizing these errors in time we can trace these back, for example, to overestimations of seasonal-annual periodicities of ecosystem respiration during spring greenup and autumn in both models. In the same frequencies, systematic misrepresentations of CO2 uptake severely affect the performance of LPJ, which is a consequence of the parsimonious representation of phenology. ORCHIDEE shows pronounced model-data disagreements in the high-frequency fluctuations of evapotranspiration across the four sites. We highlight the advantages that our novel methodology offers for a rigorous model evaluation compared to classical model evaluation approaches. We propose that ongoing model development will benefit from considering model-data (dis)agreements in the time-frequency domain.

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Dormancy release as influenced by duration of outdoor winter chilling in Florence (Italy) was studied under different photoperiodic and temperature treatments in collected twigs of two European (Ulmus glabra Huds. and Ulmus minor Mill.) and four Asian (Ulmus pumila L., Ulmus parvifolia Jacq., Ulmus macrocarpa Hance and Ulmus villosa Brandis) elm clones. Photoperiod had no effect on dormancy release, and there was no evidence that photoperiod affected bud burst during quiescence in the studied elm clones. Thermal time (day degrees >0 °C) to bud burst decreased in all the clones with increasing outdoor chilling. Although all the clones exhibited a rather weak dormancy, they significantly differed from each other. Dormancy was released earlier in the Asian than in the European clones, and the clones could be ranked from the U. pumila clone (very weak and short dormancy) to the U. minor clone (relatively stronger and longer dormancy), the other clones being intermediate. In all the clones except U. minor, the observed decrement in thermal time to bud burst was efficiently explained as an inverse exponential function of the number of chill days ≤5 °C received outdoor in autumn and winter. Endodormancy, as measured by the single-node cuttings test, was weak and short in all the clones. The latter result suggests that correlative inhibitions were largely responsible for preventing bud burst during winter in these elm clones.

Sammendrag

Chlorine - one of the most widespread elements on the Earth - is present in the environment as chloride ion or bound to organic substances. The main source of chloride ions is the oceans while organically bound chlorine (OCl) comes from various sources, including anthropogenic ones. Chlorinated organic compounds were long considered to be only industrial products; nevertheless, organochlorines occur plentifully in natural ecosystems. However, recent investigations in temperate and boreal forest ecosystems have shown them to be products of biodegradation of soil organic matter under participation of chlorine. It is important to understand both the inorganic and organic biogeochemical cycling of chlorine in order to understand processes in the forest ecosystem and dangers as a result of human activities, i.e. emission and deposition of anthropogenic chlorinated compounds as well as those from natural processes. The minireview presented below provides a survey of contemporary knowledge of the state of the art and a basis for investigations of formation and degradation of organochlorines and monitoring of chloride and organochlorines in forest ecosystems, which has not been carried out in the Czech Republic yet.